I am an evolutionary biologist interested in macroevolution. My research is mostly question driven rather than focused on any specific model system. I am interested in how phenotypic traits evolve, how species richness accumulates, the factors that drive diversification and the mechanistic explanations for the geographical species gradient. To tackle these questions I mostly use phylogenetic comparative methods, and have also contributed to the development of these methods.

I am active in teaching theoretical-practical workshops on the theory and use of phylogenetic comparative methods. To date I have taught workshops in Estonia, Sweden, Spain, Cuba, Chile, Mexico and China.

Brief CV

2007: Instituto de Ecología, Universidad Nacional Autónoma de México.  

2007-09: Evolutionary Biology Centre, Uppsala University, Sweden.


2009-12: Juan de la Cierva post-doctoral contract, Estación Biológica de Doñana, Sevilla, Spain.


2012-14: JAE-Doc post-doctoral contract, Estación Biológica de Doñana, Sevilla, Spain.


31Dugo-Cota, A., Castroviejo-Fisher, S., Vilà, C., Gonzalez-Voyer, A. 2015. A test of the integrated evolutionary speed hypothesis in a Neotropical amphibian radiation. Global Ecology and Biogeography. 24: 804-813.
30Tsuboi, M., Gonzalez-Voyer, A., Kolm, N. 2015. Functional coupling constrains craniofacial diversification in Lake Tanganyika cichlids. Biology Letters. doi: dx.doi.org/10.1098/rsbl.2014.1053
29Carrillo-Gavilán, A., Moreira, X., Zas, R., Gonzalez-Voyer, A., Vilà, M., Sampedro, L. 2015. Phylogenetic and biogeographical patterns in quantitative allocation to chemical defenses and defensive strategies in Palearctic and Nearctic pine trees (Subgenus Pinus). Journal of Biogeography. (10.1111/jbi.12444 )
28Tsuboi, M., Gonzalez-Voyer, A., Kolm, N. 2014. Phenotypic integration of brain size and head morphology in Lake Tanganyika cichlids. BMC Evolutionary Biology. 14: 39.
27Castroviejo-Fisher, S., Guayasamin, J. M., Gonzalez-Voyer, A., Vilà, C. 2014. Neotropical diversification seen through glassfrogs. Journal of Biogeography. 41: 66–80.
25Maklakov, A. A., Immler, S., Gonzalez-Voyer, A., Rönn, J. and Kolm, N. 2013. Brains and the city in passerine brids: re-analysis and confirmation of the original result. Biology Letters. 9: 20130859.
26Amcoff, M., Gonzalez-Voyer, A., Kolm, N. 2013. Evolution of egg-dummies in Tanganyikan cichlid fishes: the roles of parental care and sexual selection. Journal of Evolutionary Biology. 26: 2369-82.
24Romeralo M., Skiba A., Gonzalez-Voyer A., Schilde C., Lawal H., Kedziora S., Cavender J.C., Glöckner G., Urushihara H. and Schaap P. 2013. Analysis of phenotypic evolution in Dictyostelia highlights developmental plasticity as a likely consequence of colonial multicellularity. Proceedings of the Royal Society series B. 280: 20130976.
21vonHardenberg, A.* and Gonzalez-Voyer, A.* 2013. Disentangling evolutionary cause-effect relationships with phylogenetic confirmatory path analysis. Evolution. 67: 378–387
23Santos-Gally, R., Gonzalez-Voyer, A. and Arroyo, J. 2013. Deconstructing heterostyly: the evolutionary role of incompatibility system, pollinators and floral architecture. Evolution. 67: 2072–2082.
22Gonzalez-Voyer, A., den Tex, R. J., Castello, A. and Leonard, J. A. 2013. Evolution of acoustic and visual signals in Asian barbets. Journal of Evolutionary Biology. 26: 647–659
20Immler, S., Gonzalez-Voyer, A. and Birkhead, T. H. 2012. Distinct evolutionary patterns of morphometric sperm traits in passerine birds. Proceedings of the Royal Society B. 279 : 4174-4182.
19Fitzpatrick, J. L., Almbro, M., Gonzalez-Voyer, A., Kolm, N. and Simmons, L. W. 2012. Male contest competition and the coevolution of weaponry and testes in pinnipeds. Evolution. 66 : 3595-3604.
18Fitzpatrick, J. L., Almbro, M., Gonzalez-Voyer, A., Hamada, S., Pennington, C., Scanlan, J. and Kolm, N. 2012. Sexual selection uncouples the evolution of brain and body size in pinnipeds. Journal of Evolutionary Biology. 25 : 1321-1330.
16Duarte, H., Tejedo, M., Katzenberger, M., Marangoni, F., Baldo, D., Beltrán, J. F., Martí, D. A., Richter-Boix, A. and Gonzalez-Voyer, A. 2012. Can amphibians take the heat? Vulnerability to climate warming in subtropical and temperate larval amphibian communities. Global Change Biology. 18: 412-421.
17Cruz, F., Brennan, A. C.*, Gonzalez-Voyer, A.*, Muñoz-Fuentes, V.*, Eaaswarkhanth, M.*, Roques, S.* and Picó, X. 2012. Genetics and Genomics in Wildlife Studies : Implications for Ecology, Evolution and Conservation Biology. BioEssays. 34 : 245-246.(*Equal Contribution)
15Tsuboi, M., Gonzalez-Voyer, A., Höglund, J. and Kolm, N. 2012. Ecology and mating competition influence sexual dimorphism in Tanganyikan cichlids. Evolutionary Ecology. 26: 171-185.
14Gonzalez-Voyer, A. and Kolm, N. 2011. Rates of phenotypic evolution of ecological characters and sexual traits during the Tanganyikan cichlid adaptive radiation. Journal of Evolutionary Biology. 24: 2378–2388.
13Maklakov, A. A., Immler, S., Gonzalez-Voyer, A., Rönn, J. and Kolm, N. 2011. Brains and the city: big-brained passerine birds succeed in urban environments. Biology Letters. 7: 730-732.
12Gonzalez-Voyer, A.*, Padial, J. M.*, Castroviejo-Fisher, S., De la Riva, I. and Vilà, C. 2011. Correlates of species richness in the largest Neotropical amphibian radiation. Journal of Evolutionary Biology. 24: 931-942. (*Equal Contribution)
11Gonzalez-Voyer, A. and N. Kolm. 2010. Sex, ecology and the brain: Evolutionary correlates of brain structure volumes in Tanganyikan cichlids. PLoS ONE. 5(12), e14355.
10Kolm, N., Gonzalez-Voyer, A., Brelin, D. and Winberg, S. 2009. Evidence for small scale variation in the vertebrate brain: mating strategy and sex affect brain size and structure in wild brown trout (Salmo trutta). Journal of Evolutionary Biology. 22: 2524-2531.
6Gonzalez-Voyer, A. Fitzpatrick, J. L. and Kolm, N. 2008. Sexual selection determines parental care patterns in cichlid fishes. Evolution. 62: 2015-2026.
5Gonzalez-Voyer, A., Székely, T. and Drummond, H. 2007. Why do some siblings attack each other? Comparative analysis of broodmate aggression in birds. Evolution. 61: 1946-1951.
4 Gonzalez-Voyer, A. and Drummond, H. 2007. Is broodmate aggression really associated with direct feeding? Behaviour. 144: 373-392.
3Gonzalez-Voyer, A., Smith, K. G. and Festa-Bianchet, M. 2003. Dynamics of hunted and unhunted mountain goat Oreamnos americanus populations. Wildlife Biology. 9: 213-218.